Gha is a voiced aspirate, breathy voiced, and it fills the fourth slot of the velar class. Every one of the five classes has such a letter, and the four features that define the grid, voicing and aspiration in their four combinations, are complete once it is in place. The fifth slot of each class is the nasal, which is a different kind of sound and closes the group.
To an ear trained on a language without the series, the difference between this letter and the plain voiced stop is very slight. To Sanskrit it is the difference between one word and another, and the language keeps the distinction rigorously. Siddham gives the sound a character of its own rather than writing it as the voiced stop with an added mark, which keeps the fourfold structure of each class legible at a glance and is one reason the Indic tables were copied rather than adapted when they reached East Asia.
In frequency the letter is modest. Its main occurrences are in a limited set of roots and in the words derived from them, and a page may go by without it. What it contributes is structural: it is the letter that makes each class a complete crossing of two features rather than a partial one.
The letter also illustrates something about how the table was received. A tradition copying a syllabary for liturgical use had no obvious reason to keep a sound its own speakers could not produce, and Japanese has nothing resembling a voiced aspirate. It was kept anyway, with a settled Japanese reading attached, because what was being transmitted was the table and not a working alphabet.