Epsilon is the fifth Greek letter, from Phoenician he; its value is 5. Its later name 'e psilon' (bare e) distinguishes it from the diphthong 'ai'. Epsilon has a genuinely attested symbolic history: a mysterious letter E was displayed at the temple of Apollo at Delphi, and Plutarch devoted an essay, 'On the E at Delphi', to competing explanations of its meaning.
The Greek letters doubled as numerals in the Milesian (alphabetic) system, in which each letter carried a fixed value; the practice of summing a word's letters into a number, called isopsephy, is the Greek counterpart of Hebrew gematria and appears in inscriptions, the Greek Magical Papyri and early Christian texts. Epsilon's Phoenician ancestry places it in a borrowing whose exact point of contact, somewhere among Al Mina, Cyprus, Crete and Euboea in the Aegean world, is still disputed, though the borrowing itself, conventionally dated to around the 8th century BCE, is agreed to be the ancestor of every European alphabet still in use, including the Latin one covered elsewhere in this atlas.
The Greeks' own contribution was to give vowels dedicated letters of their own, something the consonant-only Phoenician script never did. Its isopsephy value of 5 belongs to the Milesian alphabetic numeral system, which also needed three further, otherwise obsolete letters, digamma (also called stigma), koppa and sampi, to cover every value from 1 up to 900; none of the three survives among the 24 letters in ordinary use today.